What was one reason why the Greek city-states in Persian territory were unhappy with Persian rule?

They preferred Athenian rule.

They didn't like being forced to serve in the Persian military.

They felt that the tax system was unfair.

They wanted equal rights for men and women.

There were of course multiple reasons why did were unhappy with Persian rule. But one thing that surely bothered them greatly was the fact that they were drafted into the Persian army and had to fight fellow greeks, which is obviously not a thing they would have wanted to do.